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So not only do we have to deal with non-vertical events where entire organisms are engulfed, but there are also all the lateral gene transfer events which dont fit the model of the tree. Scientists acknowledge lateral gene transfer as a phenomenon, and then we draw our trees anyway. So I think the next paradigm shift will be figuring out how to capture the combination of lateral and vertical events on depictions of the tree of life.

What are the big milestones in eukaryotic microbial evolution and can we begin to confidently place them on the tree. I think weve made surprisingly little progress in understanding diversification events at the origin of eukaryotes.